1. Department of Gastroenterology and Metabolism, Applied Life Science, Institute of Biomedical and Health Science; Hiroshima University; Hiroshima Japan
2. Liver Research Project Center; Hiroshima University; Hiroshima Japan
3. Natural Science Center for Basic Research and Development; Hiroshima University; Hiroshima Japan
4. Laboratory for Digestive Diseases, Center for Genomic Medicine; The Institute of Physical and Chemical Research (RIKEN); Hiroshima Japan