1. Laboratory of Tropical Forest Resources & Environments; Graduate School of Agriculture; Kyoto University; Kitashirakawa Oiwake-cho, Sakyo-ku; Kyoto; 606-8502; Japan
2. Forestry and Forest Products Research Institute; Matsunosato 1, Tsukuba; Ibaraki; 305-8687; Japan
3. Tropical Rain Forest Research Centre; Mulawarman University; Kampus Gunung Kelua, Jl. Kihajar Dewantara; Samarinda; 75123; East Kalimantan; Indonesia
4. Centre for Ecological Research; Kyoto University; 2-509-3 Hirano, Otsu; Shiga; 520-2113; Japan