1. Department of Botany, Marsh Life Science Building, University of Vermont, Burlington, VT05405, USA;
2. Department of Ecology and Evolutionary Biology, University of Tennessee, 569 Dabney Hall, Knoxville, TN 37996‐1610, USA;
3. Department of Biological Sciences, 202 LSB, Louisiana State University, Baton Rouge, LA 70803, USA;