1. Botany Department; University of Otago; PO Box 56; Dunedin; New Zealand
2. Department of Biology; University of North Carolina; Chapel Hill; NC; 27599-3280; USA
3. Biodiversity, Evolution and Ecology of Plants; Biocentre Klein Flottbek and Botanical Garden; University of Hamburg; Ohnhorststr. 18; 22609; Hamburg; Germany
4. Institute of Ecology and Earth Sciences; University of Tartu; Lai 40; Tartu; 51005; Estonia