1. Graduate School of Science and Technology, Niigata University, Niigata 950–2181, Japan
2. Center for Ecological Research, Kyoto University, Otsu 520–2113, Japan
3. Faculty of Agriculture, Niigata University, Niigata 950–2181, Japan
4. Tohoku Research Center, Forestry and Forest Products Research Institute, Morioka 020–0123, Japan
5. Faculty of Bio‐resource Science, Akita Prefectural University, Akita 010–0195, Japan
6. Research Institute for Humanity and Nature, Kyoto 606–8502, Japan