1. Laboratoire Evolution et Diversité Biologique, CNRS—Université Paul Sabatier, Bâtiment 4R3, 118 route de Narbonne, 31062 Toulouse, France
2. Present address: AZTI—Tecnalia/Marine Research Division, Herrera kaia portualdea z/g 20110 Pasaia (Gipuzkoa), Basque country, Spain
3. Center for Tropical Forest Science, Smithsonian Tropical Research Institute, Unit 0948, APO AA 34002‐0948, USA