1. Laboratory of Quantitative Vegetation Ecology, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China
2. Swiss Federal Institute for Forest, Snow and Landscape Research, Zürcherstrasse 111, CH‐8903 Birmensdorf, Switzerland; E‐mail
3. University of Applied Sciences Wädenswil (HSW), Centre of Expertise for Life Sciences and Facility Management, Grüental, Postfach 335, CH‐8820 Wädenswil, Switzerland; E‐mail
4. Institute of Systematic Botany, University of Zürich, Zollikerstrasse 107, CH‐8008 Zürich, Switzerland; E‐mail