1. Estación Experimental de Zonas Áridas, CSIC, General Segura 1, 04001 Almería, Spain
2. Institute of Botany of the Georgian Academy of Sciences, Kojori Road 1, Tbilisi 380007, Georgia (Republic) E‐mail
3. E‐mail
4. Division of Biological Sciences, University of Montana, Missoula, MT, 59812‐1002, USA; E‐mail