1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization and Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province Chengdu Institute of Biology, Chinese Academy of Sciences Chengdu China
2. University of Chinese Academy of Sciences Beijing China
3. Department of Zoology, College of Life Sciences Henan Normal University Xinxiang China
4. Institute of Herpetology and Liaoning Key Lab of Evolution and Biodiversity Shenyang Normal University Shenyang China