1. Center for Drug Research and Development Guangdong Pharmaceutical University Guangzhou 510006 China
2. State Key Laboratory of Applied Microbiology Southern China Guangdong Academy of Sciences Guangzhou 510070 China
3. Guangdong Provincial Key Laboratory of Microbial Culture Collection and Application, Guangdong Open Laboratory of Applied Microbiology, Guangdong Institute of Microbiology Guangdong Academy of Sciences Guangzhou 510070 China