1. Key Laboratory of Tropical Marine Bio-resources and Ecology of Chinese Academy of Sciences (LMB), Guangdong Provincial Key Laboratory of Applied Marine Biology (LAMB), South China Sea Inst. of Oceanology; Chinese Academy of Sciences; No. 164, West Xingang Rd. Guangzhou Guangdong 510301 PR China
2. Univ. of Chinese Academy of Sciences; Beijing 100049 PR China