1. Dept of Zoology; Faculty of Biology; Seville Spain
2. Environment Canada, Saskatoon, Saskatchewan, Canada, and Univ. of Western Ontario; London ON Canada
3. Dept of Anatomy; Cellular Biology and Zoology, Univ. of Extremadura; Badajoz Spain
4. Laboratoire dfEcologie, Systématique et Evolution, CNRS UMR 8079, Univ. Paris-Sud; Orsay Cedex France