1. School of Geography, Archaeology and Environmental Studies; University of the Witwatersrand; Johannesburg South Africa
2. Department of Anatomy; University of Pretoria; Pretoria South Africa
3. Laboratoire d'Anthropologie Moléculaire et Imagerie de Synthèse; UMR 5288 CNRS-Université de Toulouse (Paul Sabatier); Toulouse Cedex France
4. Institut de Recherche en Informatique de Toulouse; UMR 5505 CNRS-Université de Toulouse (Paul Sabatier); Toulouse Cedex France
5. Radiation Science Department; South African Nuclear Energy Corporation (Necsa); Pelindaba South Africa
6. Institut du Cerveau et de la Moelle épinière; Aramis Team; INRIA Paris; Sorbonne Universités; UPMC Université Paris 06 UMR S 1127; Inserm U 1127, CNRS UMR 7225; Paris France
7. Department of African Zoology; Royal Museum for Central Africa; Tervuren Belgium
8. Laboratory of Histology and Neuropathology; Université Libre de Bruxelles; Brussels Belgium
9. Department of Anatomy and Histology; Sefako Makgatho Health Sciences University; Pretoria South Africa
10. Montpellier Laboratory of Informatics; Robotics and Microelectronics; UMR 5506 CNRS; Université de Montpellier; Montpellier France
11. Evolutionary Studies Institute and School of Geosciences; University of the Witwatersrand; Johannesburg South Africa