1. Hainan Aquaculture Breeding Engineering Research Center College of Marine Sciences Hainan University Haikou Hainan570228 China
2. Department of Marine Sciences University of Connecticut Groton Connecticut06340 USA
3. State Key Laboratory of Marine Environmental Science Xiamen University Xiamen Fujian361102 China
4. Key Laboratory of Coastal Zone Environmental Processes Yantai Institute of Coastal Zone Research Chinese Academy of Sciences Yantai Shandong264003 China