1. NLA University College; PO Box 74 Sandviken N-5812 Bergen Norway
2. Department of Biology; University of Bergen; PO Box 7803 N-5020 Bergen Norway
3. Systematics and Evolution Group, Museum of Natural History and Archaeology; Section of Natural History; Norwegian University of Science and Technology; N-7491 Trondheim Norway
4. Department of Geography; University of Bergen; PO Box 7802 N-5020 Bergen Norway