1. School of Earth, Atmospheric & Environmental Sciences; The University of Manchester; Manchester M13 9PL UK
2. British Antarctic Survey; Natural Environment Research Council; Cambridge CB3 0ET UK
3. Department of Arctic Biology; the University Centre in Svalbard; P.O. Box 156 N-9171 Longyearbyen Svalbard
4. Institute of Bio- and Geosciences; Agrosphere (IBG-3); Forschungszentrum Jülich GmbH; Wilhelm-Johnen-Straße 52425 Jülich Germany
5. Sustainable Soils and Grassland Systems Department; Rothamsted Research; North Wyke Okehampton Devon EX20 2SB UK