1. Institute of Marine Biology; National Taiwan Ocean University; Keelung 20224 Taiwan
2. Simon F. S. Li Marine Science Laboratory; School of Life Sciences; The Chinese University of Hong Kong; Shatin NT Hong Kong China
3. Jiangsu Key Laboratory of Marine Biotechnology/College of Marine Science; Huaihai Institute of Technology; Lianyungang 222005 China
4. Department of Science and Environmental Studies; The Education University of Hong Kong; Taipo Hong Kong China
5. Biodiversity Research Center; Academia Sinica; Taipei 115 Taiwan