1. Hainan Jianfengling Forest Ecosystem National Field Science Observation and Research Station, Research Institute of Tropical Forestry Chinese Academy of Forestry Guangzhou 510520 China
2. College of Life Sciences Gannan Normal University Ganzhou 341000 Jiangxi China
3. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
4. Departamento de Zoología, Facultad de Biología Universidad de Santiago de Compostela 15782 Santiago de Compostela Spain
5. Hainan Academy of Forest (Hainan Academy of Mangrove) Haikou 571100 China