1. CoMPLEX, Centre for Mathematics and Physics in the Life Sciences and Experimental Biology University College London London United Kingdom
2. Department of Genetics Evolution and Environment, University, College London London United Kingdom
3. Environment and Sustainability Institute University of Exeter, Penryn, Campus Tremough United Kingdom
4. Department of Mathematics University of Surrey Guildford GU2 7XH United Kingdom
5. Mathematical Sciences University of Southampton Highfield Southampton SO17 1BJ United Kingdom