1. Inst. for Evolution and Ecology, Dept of Biology, Univ. of Tübingen Auf der Morgenstelle 5 DE‐72076 Tübingen Germany
2. Biogeophysics, Inst. of Soil Science and Land Evaluation, Univ. of Hohenheim Stuttgart Germany
3. Dept of Environmental Biology, The School of Environmental Sciences, Univ. of Guelph Guelph ON N1G 2W1 Canada