1. College of Agriculture and Biotechnology; Zhejiang University; Hangzhou 310058 China
2. State Key Laboratory Breeding Base for Zhejiang Sustainable Pest and Disease Control; Ministry of Agriculture Key Laboratory of Biotechnology in Plant Protection; Zhejiang Provincial Key Laboratory of Plant Virology; Institute of Virology and Biotechnology; Zhejiang Academy of Agricultural Sciences; Hangzhou 310021 China
3. Taizhou Academy of Agricultural Sciences; Linhai 317000 China
4. Zhejiang Provincial Institute for the Control of Agrochemicals; Hangzhou 310020 China