1. Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering; School of Marine Sciences; Sun Yat-sen University; Guangzhou China
2. State Key Laboratory of Biocontrol/; School of Marine Sciences; Sun Yat-sen University; Guangzhou China
3. Institute of Aquatic Economic Animals, and Guangdong Province Key Laboratory for Aquatic Economic Animals; Sun Yat-sen University; Guangzhou China