1. Department of Biology; University of York; Heslington York YO10 5DD UK
2. York Institute for Tropical Ecosystems; Environment Department; University of York; Heslington York YO10 5NG UK
3. Institute of Biodiversity, Animal Health and Comparative Medicine; University of Glasgow; Glasgow G12 8QQ UK
4. Centre for African Ecology; School of Animal, Plant and Environmental Sciences; University of the Witwatersrand; Private Bag Johannesburg South Africa
5. Natural Resources and the Environment; CSIR; PO Box 395 Pretoria 0001 South Africa
6. Department of Biology; Wake Forest University; 049 Winston Hall Winston-Salem North Carolina 27106 USA
7. Ecology and Evolutionary Biology; Princeton University; Eno Hall Princeton NJ 08540 USA
8. South African Environmental Observation Network (SAEON); Ndlovu Node; Private Bag x1021 Phalaborwa Kruger National Park 1390 South Africa
9. Department of Earth, Ocean & Ecological Sciences; University of Liverpool; Liverpool L69 3GP UK
10. Department of Zoology & Entomology; University of Pretoria; Private Bag X20 Pretoria 0028 South Africa