1. Fisheries Department, Faculty of Natural Resources University of Guilan Sowmeh Sara Iran
2. Department of Fisheries, Hamoun International Wetland Research Institute University of Zabol Zabol Iran
3. International Sturgeon Research Institute Agricultural Research Education and Extension Organization Rasht Iran
4. Cell Biology in Environmental Toxicology Research Group, Department of Zoology and Animal Cell Biology, Faculty of Science and Technology and Research Centre for Experimental Marine Biology and Biotechnology University of the Basque Country Basque Country Spain
5. Department of Zoology University of Otago Dunedin New Zealand