1. Museum of Zoology, Graduate Program in Systematics, Animal Taxonomy and Biodiversity University of São Paulo São Paulo Brazil
2. Current address: Department of Genetics and Evolutionary Biology, Institute of Biosciences University of São Paulo São Paulo Brazil
3. Department of Genetics and Evolutionary Biology, Institute of Biosciences University of São Paulo São Paulo Brazil
4. Department of Entomology and Acarology, Luiz de Queiroz College of Agriculture (ESALQ), Graduate Program in Entomology University of São Paulo Piracicaba Brazil
5. Department of Ecology and Evolutionary Biology, Institute of Environmental, Chemical and Pharmaceutical Sciences Federal University of São Paulo Diadema Brazil