1. Key Laboratory of Alpine Ecology and Biodiversity; Institute of Tibetan Plateau Research; Chinese Academy of Sciences; 16 Lincui Road Chaoyang District 100101 Beijing China
2. CAS Center for Excellence in Tibetan Plateau Earth Sciences; 16 Lincui Road, Chaoyang District, 100101 Beijing China
3. Department of Biology, Centre of Excellence PLECO (Plant and Vegetation Ecology); University of Antwerp, Universiteitsplein 1; B-2610 Wilrijk Belgium