1. Department of Biological Sciences; Macquarie University; North Ryde NSW 2109 Australia
2. Hawkesbury Institute for the Environment; University of Western Sydney; Locked Bag 1797 Penrith NSW 2751 Australia
3. Geography Department; College of Life and Environmental Sciences; University of Exeter; Exeter EX4 4RJ UK
4. Centre for Ecology and Hydrology; Wallingford OX10 8BB UK
5. Biogeochemical Integration Department; Max Planck Institute for Biogeochemistry; Hans-Knöll-Str. 10 07745 Jena Germany