1. Marine Biological Association of the United Kingdom, The Laboratory; Citadel Hill Plymouth PL1 2PB U.K.
2. Ocean and Earth Science, National Oceanography Centre Southampton; University of Southampton; Waterfront Campus, European Way Southampton SO14 3ZH U.K.
3. Centre for Biological Sciences; Building 85, University of Southampton, Highfield Campus Southampton SO17 1BJ U.K.