1. Red Sea Research Center, Division of Biological and Environmental Science and Engineering King Abdullah University of Science and Technology Thuwal Saudi Arabia
2. Center for Environment and Water, Research Institute King Fahd University of Petroleum and Minerals Dhahran Saudi Arabia
3. Department of Marine Biology and Environmental Sciences, Faculty of Agriculture University of Miyazaki Miyazaki Japan
4. Division of Physical Science and Engineering King Abdullah University of Science and Technology Thuwal Saudi Arabia