1. Department of Biological and Environmental Sciences; Göteborg University; Box 461 Göteborg SE-405 30 Sweden
2. National Oceanic and Atmospheric Administration; National Marine Fisheries Service; Northeast Fisheries Science Center; Milford Laboratory; 212 Rogers Avenue Milford Connecticut 06460 USA
3. Hollings Marine Laboratory; College of Charleston; 331 Fort Johnson Rd Charleston South Carolina 29412 USA