1. Department of Ecology & Evolutionary Biology, University of Colorado, Boulder, CO 80309‐0334, USA;
2. Present address: Instituto de Investigaciones Agropecuarias, Unidad de Biotecnología, INIA Carillanca, Casilla 58‐D, Temuco, Chile
3. Department of Plant Biology, 256 Plant Science Building, Cornell University, Ithaca, NY 14853, USA;
4. Biology Department, Bowdoin College, Brunswick, ME 04011, USA;