1. Institute of Biological, Environmental and Rural Sciences, Aberystwyth, Wales SY23 3DA, UK
2. Department of Agriculture, University of Zaragoza, High Polytechnic School of Huesca, Ctra. Cuarte km 1, ES–22071 Huesca, Spain
3. Department of Plant Anatomy and Cytology, Faculty of Biology and Environmental Protection, University of Silesia, PL–40‐032 Katowice, Poland
4. USDA ARS Western Regional Research Center, Albany, CA 94710 USA