1. W.K. Kellogg Biological Station Michigan State University, 3700 East Gull Lake Drive, Hickory Corners, MI 49060, USA
2. Department of Plant Biology, Michigan State University, 166 Plant Biology Building, East Lansing, MI 48824, USA
3. Department of Microbiology and Molecular Genetics, Michigan State University, 2215 Biomedical Physical Sciences, East Lansing, MI 48824, USA