1. Institut de Botanique, Université de Liege, B‐22 Sart Tilman, B‐4000 Liege, Belgium
2. Biology Department, Duke University, Box 90338 Durham, NC 27708, USA
3. National Herbarium of New South Wales, Royal Botanic Gardens Sydney, Mrs Macquaries Road, Sydney, NSW 2000, Australia
4. Dept. Systématique et Evolution, Muséum National d’Histoire Naturelle, 57 rue Cuvier, 75231 Paris cedex 05, France