1. Department of Biology, University of Virginia, Charlottesville, VA 22904, USA
2. Rosenstiel School of Marine and Atmospheric Science, University of Miami, Miami, FL 33149‐1098, USA
3. Department of Biology, University of Miami, Coral Gables, FL 33124‐0421, USA
4. School of Marine and Atmospheric Sciences, Stony Brook University, Stony Brook, NY 11794‐5000, USA
5. Department of Environmental Sciences, University of Virginia, Charlottesville, VA 22904‐4123, USA
6. Xishuangbanna Tropical Botanic Garden, Melung, Xishuangbanna, Yunnan, China