1. Key Laboratory of Biodiversity and Biogeography, Kunming Institute of Botany, Chinese Academy of Sciences, Lanhei Road 132, Heilongtan, Kunming 650204, Yunnan, China
2. Graduate University of the Chinese Academy of Sciences, Beijing 100049, China
3. Biology Department, Dartmouth College, Hanover, NH 03755, USA
4. Institute of Tibetan Plateau Research at Kunming, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650204, China
5. Sichuan Academy of Forestry, Chengdu 610081, China