1. Department of Biology, University of Missouri – St. Louis, Research 223, One University Boulevard, St. Louis, MO 63121, USA
2. Department of Ecology and Evolutionary Biology, The University of Kansas, 8009 Haworth Hall, 1200 Sunnyside Avenue, Lawrence, KS 66045, USA
3. Laboratory of Genetics, University of Wisconsin, Madison, WI 53706, USA