1. Department of Botany, University of British Columbia, 6270 University BLVD, Vancouver, BC V6T 1Z4, Canada;
2. Cirad‐CA, Station Roujol, 97170 Petit‐Bourg, Guadeloupe, French West Indies;
3. Department of Geography, Indiana University, 701 E. Kirkwood Avenue, Bloomington, IN 47405, USA