1. Department of Forest Ecology and Management, SLU, SE‐901 83 Umeå, Sweden;
2. Department of Plant Physiology and Umeå Plant Science Centre, University of Umeå, SE‐901 87 Umeå, Sweden;
3. Department of Forest and Soil Science, University of Natural Resources and Applied Life Sciences, BOKU, Peter Jordan‐Strasse 82, A‐1190 Vienna, Austria;
4. Unit for Field‐based Forest Research, SLU, Vindeln Experimental Forests and Svartberget Research Station, SLU, SE‐922 91 Vindeln, Sweden;
5. Southern Swedish Forest Research Centre, SLU, PO Box 49, SE‐230 53 Alnarp, Sweden;
6. Department of Forest Genetics and Plant Physiology and Umeå Plant Science Centre, SLU, SE‐901 87 Umeå, Sweden