1. School of Biological Sciences, Plant Sciences Laboratories, University of Reading, Reading, Berkshire RG6 6AS, UK;
2. CSIRO Entomology and CRC for Australian Weed Management, Long Pocket Laboratories, 120 Meiers Road, Indooroopilly, QLD 4068, Australia;
3. Department of Plant Sciences, University of Oxford, South Parks Road, Oxford OX1 3RB, UK