1. Department of Botany, University of Guelph, Guelph, Ontario N1G 2W1, Canada;
2. Present address: Department of Ecology & Evolutionary Biology, University of Toronto, 25 Willcocks Street, Toronto, Ontario M5S 3B2, Canada
3. School of Biological Sciences, Washington State University, PO Box 644236, Pullman, WA 99164‐4236, USA;