1. Department of Organismic and Evolutionary Biology, Harvard University, 22 Divinity Ave., Cambridge, MA 02138, USA;
2. Department of Plant Sciences and Plant Pathology, Montana State University, PO Box 173150, Bozeman, MT 59717‐3150, USA;
3. Institute of Genetics, Adrian Building, University of Leicester, University Road, Leicester LE1 7RH, UK