1. Department of Biological Sciences, Stanford University, Stanford, CA 94305, USA;
2. (Present address) Department of Integrative Biology, University of California Berkeley, Berkeley, CA 94720, USA
3. Instituto de Ecologia, Universidad Nacional Autonoma de Mexico, Apartado Postal 70‐275, UNAM, 04510 Mexico, D.F.;
4. Department of Integrative Biology, University of California Berkeley, Berkeley, CA 94720, USA;