1. Department of Soil Environmental Sciences, Research Center for Eco‐environmental Sciences, Chinese Academy of Sciences, 18 Shuangqing RD, Beijing 100085, People's Republic of China;
2. Department of Biochemistry and Molecular Biology, Wayne State University, School of Medicine, Detroit, MI 48201, USA;
3. Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing 100083, China