1. Robert W. Holley Center for Agriculture and Health, USDA‐ARS, Cornell University, Ithaca, NY 14853, USA
2. Department of Plant Breeding and Genetics, Cornell University, Ithaca, NY 14853, USA
3. The State Key Laboratory of Pharmaceutical Biotechnology, School of Life Sciences, Nanjing University, Nanjing 210093, China
4. The State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing 100101, China