1. Division of Plant Pathology and Microbiology, School of Plant Sciences, The University of Arizona, Tucson, AZ 85721, USA
2. Department of Biological Sciences, Northern Arizona University, Flagstaff, AZ 86011, USA
3. Imperial College London and Royal Botanic Gardens, Kew TW9 3DS, UK
4. Department of Biological Sciences, Barnard College, Columbia University, New York, NY 10027, USA