1. Institute of Biology, Martin‐Luther‐University Halle‐Wittenberg, 06099 Halle (Saale), Germany
2. Present address: Institute of Genetics, University of Munich (LMU), Großhaderner Straße 2, 82152 Martinsried, Germany
3. The Sainsbury Laboratory, Norwich, NR4 7UH, UK
4. National Key Laboratory of Crop Genetic Improvement, National Center of Plant Gene Research (Wuhan), Huazhong Agricultural University, Wuhan 430070, PR China