1. College of Forest Resources, University of Washington, Box 352100, Seattle, WA 98195‐2100, USA;
2. Present address: Systematic Botany and Mycology Lab, USDA ARS Room 304, BLD 011A, 10300 Baltimore Avenue, Bettsville, MD 20705‐2350, USA
3. Biology Department, University of Washington, Box 355325, Seattle, WA 98195, USA;