1. Department of Horticulture, Pennsylvania State University, University Park, PA 16802, USA;
2. Intercollege Graduate Programs in Plant Physiology and Ecology, Pennsylvania State University, University Park, PA 16802, USA;
3. Current address: USDA‐ARS Pasture Systems and Watershed Management Research Unit, University Park, PA 16802, USA;
4. Current address: Department of Botany/Microbiology, Ohio Wesleyan University, Delaware, OH 43015, USA
5. Department of Horticultural Sciences, New York State Agricultural Experiment Station, Cornell University, Fredonia, NY 14063, USA;
6. Department of Horticultural Sciences, New York State Agricultural Experiment Station, Cornell University, Geneva, NY 14456, USA;