1. School of Biological Sciences, Victoria University of Wellington, PO Box 600, Wellington, New Zealand
2. Department of Chemistry, The New Zealand Institute for Plant & Food Research Limited, University of Otago, PO Box 56, Dunedin, New Zealand
3. Department of Biology, High Point University, University Station 3591, High Point, NC 27262, USA
4. Department of Evolutionary Biology and Animal Ecology, Faculty of Biology, University of Freiburg, Hauptstrasse 1, D‐79104 Freiburg, Germany